a foe of the X-Men and the Avengers. She has also been known as Viper and Madame Hydra at various points in her history. Since her original introduction...

Valentina Allegra de Fontaine succeeded Viper as Madame Hydra. She wears an elaborate tentacle headdress and elaborate HYDRA robes. Elisa Sinclair has appeared...

main heads of Hydra: the Viper, Madame Hydra, Kraken, and the Hive, as well as resurrecting the Gorgon for the purpose of showing Hydra's "true self" to...

to have succeeded Viper as Madame Hydra. The Supreme Hydra (also called the Hydra Supreme) is a term given to the leader of Hydra. At other times, the...

Wolverine's prominent enemies include Sabretooth, the Hellfire Club, Viper (Madame Hydra), and Lady Deathstrike. His first love was Rose O'Hara, a close friend...

Madame Hydra, hybridized with Viper, appears in The Avengers: Earth's Mightiest Heroes, voiced by Vanessa Marshall. This version is an agent of HYDRA...

the HYDRA cause, to the extent that Baron Strucker appointed it as a figurehead alongside himself, Viper, Gorgon, Kraken, and the new Madame HYDRA in the...

that Madame Hydra had procured from the Yashidas. Madame Hydra presents the Japanese box to Leviathan leader Magadan, much to the dismay of Viper. However...

all Hydra leaders (Kraken, Viper, Madame Hydra, and Hive), and resurrects the Gorgon for the purpose of "letting the world become aware of Hydra's true...

original Viper, his brother the original Eel, and the Cobra. The second Squad consisted of the second Viper (who was also formerly Madame Hydra), the Eel...