The Bulldog Shale is a formation of Early Cretaceous age (Aptian to Albian stages) that forms part of the Marree Subgroup of the Rolling Downs Group , located in the Eromanga Basin of South Australia , Queensland and New South Wales .[ 1] [ 2]
It is the lowermost unit in the Marree Subgroup, overlying the Cadna-owie Formation and is overlain by the Coorikiana Sandstone . The formation dates to the Aptian to Albian stages of the Early Cretaceous.[ 3] The Bulldog Shale is composed of finely laminated carbonaceous and pyritic mudstone and claystone beds. Weathering has caused heavy leaching and bleaching in some regions of the Bulldog Shale, including those around Coober Pedy, so that the rocks are white or multicolored. These horizons contain rich opal deposits. Horizons without this bleaching are primarily composed of organic-rich shale . Gypsum , in addition to carbonate limestone concretions rich in fossils are common in these unbleached shaly horizons.[ 4]
The Bulldog Shale has yielded fossils of plants , invertebrates , fish , and reptiles .[ 5] The macroinvertebrate fauna of this formation includes several molluscs , such belemnites , gastropods , and bivalves . Fish are represented by chimaeras [ 6] and ray-finned fish (these include teleosts )[ 6] and a lungfish . Sharks are conspicuously absent in the Bulldog Shale.[ 4] Many plesiosaurs are known from the formation, including leptocleidids , elasmosaurids ,[ 4] pliosaurids , and possible polycotylids . Ichthyosaurs are also present.[ 7] Archosaur fossils from the Bulldog Shale are rare, and are represented mostly indeterminate specimens, some of which can be assigned to Dinosauria .[ 8] Due to the coastal location of the Bulldog Shale, large amounts of wood have also been recovered in this formation.[ 4]
Plesiosaurs reported from the Bulldog Shale Genus Species Location Material Notes Images Kronosaurus K. queenslandicus Teeth Originally referred to cf. K. sp.[ 4] [ 7] Leptocleidus L. sp. Reclassified as Umoonasaurus [ 7] Opallionectes O. andamookaensis Lunatic Hill opal field Opalized incomplete articulated skeleton A plesiosaur of uncertain classification[ 4] [ 7] Umoonasaurus U. demoscyllus Zorba Extension Opal Field, Andamooka opal fields, Curdimurka area, Neales River region Opalized skulls and skeletons A small leptocleidid[ 9] plesiosaur with three crests on its head[ 2] Elasmosauridae Indet. Andamooka Partial skeletons and several fragments [ 4] Polycotylidae Indet. Hermit Hill Fragmentary specimen [ 4]
Ichthyosaurs reported from the Bulldog Shale Genus Species Location Material Notes Images Platypterygius P. sp. Bopeechee Siding Fragmentary cranial and postcranial material The specimen SAM P14508 shows evidence of healed bite marks.[ 5]
Color key Notes Uncertain or tentative taxa are in small text ; crossed out taxa are discredited.
^ Geoscience Australia. "Stratigraphic Unit Details: Bulldog Shale" . Australian Stratigraphic Units Database . Retrieved 26 June 2020 . ^ a b Kear, Benjamin P.; Schroeder, Natalie I.; Lee, Michael S.Y. (2006). "An archaic crested plesiosaur in opal from the Lower Cretaceous high-latitude deposits of Australia" . Biology Letters . 2 (4): 615–619. doi :10.1098/rsbl.2006.0504 . PMC 1833998 . PMID 17148303 . ^ Bulldog Shale at Fossilworks .org ^ a b c d e f g h Kear, Benjamin P. (2006). "Marine reptiles from the Lower Cretaceous of South Australia: elements of a high-latitude cold-water assemblage" . Palaeontology . 49 (4): 837–856. Bibcode :2006Palgy..49..837K . doi :10.1111/j.1475-4983.2006.00569.x . S2CID 128232205 . ^ a b Zammit, Maria; Kear, Benjamin P. (2011). "Healed bite marks on a Cretaceous ichthyosaur" (PDF) . Acta Palaeontologica Polonica . 56 (4): 859–863. doi :10.4202/app.2010.0117 . ^ a b McHenry, Colin R. (2009). Devourer of Gods: The palaeoecology of the Cretaceous pliosaur Kronosaurus queenslandicus (Thesis). The University of Newcastle. ^ a b c d Kear, Benjamin P. (2016). "Cretaceous marine amniotes of Australia: perspectives on a decade of new research" (PDF) . Memoirs of Museum Victoria . 74 : 17–28. doi :10.24199/j.mmv.2016.74.03 . ^ a b c Barrett, Paul M.; Kear, Benjamin P.; Benson, Roger B.J. (2010). "Opalized archosaur remains from the Bulldog Shale (Aptian: Lower Cretaceous) of South Australia" (PDF) . Alcheringa . 34 (3): 1–9. Bibcode :2010Alch...34..293B . doi :10.1080/03115511003664440 . ISSN 0311-5518 . ^ Parrilla-Bel, Jara; Canudo, José Ignacio (2015). "On the presence of plesiosaurs in the Blesa Formation (Barremian) in Teruel (Spain)" . Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen . 278 (2): 213–227. doi :10.1127/njgpa/2015/0526 . ^ Kear, Benjamin P. (2011). Dinosaurs in Australia : Mesozoic life from the Southern Continent . Robert J. Hamilton-Bruce, CSIRO Publishing. Collingwood, Vic.: CSIRO Pub. p. 86. ISBN 978-0-643-10169-2 . OCLC 692219338 . ^ Popov, Evgeny V. (2020-12-10). "Systematic reassessment of Edaphodon eyrensis Long, 1985 (Holocephali, Chimaeroidei) from the Early Cretaceous of South Australia" . Journal of Vertebrate Paleontology . 40 (6): e1884564. Bibcode :2020JVPal..40E4564P . doi :10.1080/02724634.2020.1884564 . ISSN 0272-4634 . ^ a b Kear, Benjamin P. (2011). Dinosaurs in Australia : Mesozoic life from the Southern Continent . Robert J. Hamilton-Bruce, CSIRO Publishing. Collingwood, Vic.: CSIRO Pub. p. 82. ISBN 978-0-643-10169-2 . OCLC 692219338 . ^ a b Kear, Benjamin P. (2011). Dinosaurs in Australia : Mesozoic life from the Southern Continent . Robert J. Hamilton-Bruce, CSIRO Publishing. Collingwood, Vic.: CSIRO Pub. p. 83. ISBN 978-0-643-10169-2 . OCLC 692219338 . B. P. Kear. 2007. A juvenile pliosauroid plesiosaur (Reptilia: Sauropterygia) from the Lower Cretaceous of South Australia. Journal of Paleontology 81(1):154-162 R. E. Molnar. 1991. Fossil reptiles in Australia. In P. Vickers-Rich, J. M. Monaghan, R. F. Baird, & T. H. Rich (eds.), Vertebrate Paleontology of Australasia 605-702 R. E. Molnar. 1980. Australian late Mesozoic continental tetrapods: some implications. Mémoires de la Société Géologique de France, Nouvelle Série 139:131-143