Charmouth Mudstone Formation
Charmouth Mudstone Formation | |
---|---|
Stratigraphic range: Lower Sinemurian– Lower Pliensbachian Semicostatum-lowermost Margaritatus | |
Type | Formation |
Unit of | Lias Group |
Sub-units | Shales With Beef Member, Black Ven Marl Member, Belemnite Marl Member, Green Ammonite Member |
Underlies | Dyrham Formation, Marlstone Rock Formation, unconformity with Gault (Charmouth area) |
Overlies | Blue Lias Formation, Scunthorpe Mudstone |
Area | Wessex Basin, Worcester Basin, East Midland Shelf |
Thickness | ~335 m |
Lithology | |
Primary | Shale, mudstone |
Other | Limestone, sandstone |
Location | |
Region | England |
Country | United Kingdom |
Type section | |
Named for | Charmouth |
Location | Cliff and foreshore exposures between Seven Rock Point and Golden Cap, Dorset |
The Charmouth Mudstone Formation is a geological formation in England, dating to the Early Jurassic (Sinemurian–Pliensbachian).[1] It forms part of the lower Lias Group. It is most prominently exposed at its type locality in cliff section between Lyme Regis and Charmouth (alongside the underlying Blue Lias) but onshore it extends northwards to Market Weighton, Yorkshire, and in the subsurface of the East Midlands Shelf and Wessex Basin. The formation is notable for its fossils, including those of ammonites and marine reptiles and rare dinosaur remains. The formation played a prominent role in the history of early paleontology, with its Lyme Regis-Charmouth exposure being frequented by fossil collectors including Mary Anning.
Stratigraphy
[edit]Shales With Beef Member
[edit]The Shales With Beef Member is around 28–30 metres thick in the Lyme Regis-Charmouth region and predominantly consists of thinly bedded medium to dark grey mudstone, blocky calcareous pale-weathering mudstone and brown-grey organic-rich mudstones with frequent bedding parallel veins of fibrous calcite ("beef"), that are usually less than 10 centimetres thick. Several beds of nodular and tabular limestone are also present. It is the lowest unit of the formation and directly overlies the Blue Lias Formation, with the boundary being marked by a prominent bioturbated horizon. Notable persistent marker beds within the member include the laminated calcareous siltstone "Fish Bed", "Table Ledge", which consists of lens beds of limestones with mud content with nests of rhynchonellid brachiopods, the Devonshire Head and the Spittles limestones and the Birchi Nodules (which are septarian concretions) The upper boundary with the Black Ven Marl Member is marked by the prominent laterally persistent limestone Birchi Tabular Bed.[2]
Black Ven Marl Member
[edit]The Black Ven Marl Member is around 43 metres thick consists of thinly bedded dark mudstones,[3] with several laterally persistent cementstone horizons, notable horizons include the Lower and Upper Cement beds and the Stellare nodules.
Belemnite Marl Member
[edit]The Belemnite Marl Member is around 20 to 27 metres thick, and consists of interbedded pale and dark grey calcareous mudstone, with numerous belemnites, hence the name. The top of the member is marked by the Belemnite Stone Bed[4]
Green Ammonite Member
[edit]The Green Ammonite Member is up to 31 metres thick predominantly consists of medium grey mudstones, with 3 limestone horizons, Lower Limestone; Red Band, and Upper Limestone, it is conformably overlain by the Dyrham Formation in some areas,[5] but in the Charmouth area there is an erosive unconformable boundary with the much younger Early Cretaceous (Albian) aged Gault clays.
Paleobiota
[edit]Ammonites
[edit]Ammonites of the Charmouth Mudstone Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Apoderoceras | A. cf. dunrobinense, A. subtriangulare | |||||
Arnioceras | A. semicostatum | Semicostatum Zone | ||||
Asteroceras | A. obtusum | Obtusum zone | ||||
Caenisites | ||||||
Cymbites | ||||||
Echioceras | E. raricostatum | Raricostatum Zone | ||||
Eoderoceras | E. bispinigerum | |||||
Gemmellaroceras | G. cf. peregrinum | Belemnite Marls | ||||
Oxynoticeras | O. williamsi | |||||
Phricodoceras | P. taylori, P. lamellosum | Belemnite Marls | ||||
Promicroceras | P. planicosta | Obtusum Zone | ||||
Radstockiceras | R. complicatum | |||||
Tragophylloceras | T. ibex, T. loscombi | Green Ammonite Member | ||||
Uptonia | U. bronni | |||||
Xipheroceras |
Sauropterygia
[edit]Sauropterygia of the Charmouth Mudstone Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
A. rostratus | Charmouth | Arnioceras semicostatum-Echioceras raricostatum ammonoid zone | BMNH 38525 (holotype skeleton) | A marine sauropterygian, member of the family Rhomaleosauridae | ||
A. conybeari |
| Asteroceras obtusum, Arnioceras semicostatum-Echioceras raricostatum ammonoid zones |
| A marine sauropterygian, early member of the family Pliosauridae. It was first identified as Plesiosaurus conybeari. | ||
? P. sp. |
| Prodactylioceras davoei ammonoid zone |
| A marine sauropterygian, type member of the family Plesiosauroidea inside Plesiosauria. A juvenile specimen that resembles those assigned to the genus Plesiosaurus. Its assignation is dubious. | ||
P. dolichodeirus |
| Echioceras raricostatum ammonoid zone |
| A marine sauropterygian, type member of the family Plesiosauroidea inside Plesiosauria. The typical plesiosaur, and one of the most common found on the formation. | ||
Thaumatodracon[10] | T. wiedenrothi | Between Lyme Regis and Charmouth | Black Ven | NLMH 106.058, "a partial skeleton comprising a complete cranium, mandible, articulated cervical series, and indeterminate fragmentary remains" | A rhomaleosaurid |
Ichthyosauria
[edit]Ichthyosauria of the Charmouth Mudstone Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Ichthyosauria indet. | Indeterminate | Lyme Regis, Charmouth | Indeterminate remains | Un-attributable to any genus | ||
I. anningae[11] | Charmouth | Stonebarrow Marls Member | DONMG:1983.98 Subadult specimen | |||
Ichthyosaurus sp. | West of Westhay Water, Stonebarrow | Stonebarrow Marls Member | NHMUK R15907, partial skeleton[12] | Considered to belong to I. communis by Bennett et al. 2012, considered indeterminate within Ichthyosaurus by Lomax and Massare 2015 | ||
L. moorei[13] | Seatown | Belemnite Marls Member | BMNH R14370 partial anterior skeleton with most complete skull | A leptonectid ichthyosaur | ||
L. solei[14] | Seatown | Acanthopleuroceras valdani Subzone, Tragophylloceras ibex Zone, lower Pliensbachian | NMW 91.296.2.2 "a large, isolated and incomplete forefin" | |||
Temnodontosaurus | T. platyodon[15] |
Thalattosuchia
[edit]Thalattosuchia of the Charmouth Mudstone Formation | |||||||
---|---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images | |
Thalattosuchia[16] | Indeterminate |
| Black Ven Mudstone Member |
| Early Thalattosaurian specimens that can´t be referred to any concrete genus. Includes a possible early diverging metriorhynchoid | ||
T. hingleyae | 1 kilometer west of the Charmouth Heritage Coast Centre | Belemnite Marl Member | LRM 2021/45, a partial skeleton including cranial material, cervical, dorsal, and caudal vertebrae, ribs, right pectoral girdle, partial limb bones, and an osteoderm | An early diverging thalattosuchian. |
Pterosauria
[edit]Pterosauria of the Charmouth Mudstone Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
D. macronyx |
| Black Ven Marl Member |
| A basal pterosaur and the type member of the family Dimorphodontidae. | ||
Indeterminate |
| Black Ven Marl Member |
| A basal dimorphodontid pterosaur. Known as "Charmouth dimorphodontid" it shares resemblance with the genus Dimorphodon. | ||
Indeterminate[21] |
| Lower Pliensbachian member | Associated remains of the rostrum and mandibles | A basal dimorphodontid pterosaur. The relatively deep rostrum and extreme size dimorphism in the dentition show clear similarities to Dimorphodon, also from the Lias of Dorset. The distinctive dentition, in which the first four pairs of rostral teeth and two pairs of mandibular teeth are relatively large and fang-like, while the remaining teeth are remarkably small and short.[21] |
Dinosauria
[edit]Dinosauria of the Charmouth Mudstone Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
"Megalosaurus" lydekkeri | Lyme Regis | Black Ven Marl Member |
| A theropod, possibly a member of Coelophysidae inside Neotheropoda. Invalid and non diagnostic. It was identified as Zanclodon(?) sp. b., Megalosaurus lydekkeri and reassigned as Magnosaurus(?) lydekkeri.[24] Mortimer found that was changed to "Megalosaurus" woodwardi later.[24] It was found on mostly of recent works to be non diagnostic, probably a coelophysoid.[23] | ||
D. normani | Lower cliff face, Charmouth | Black Ven Marl Member (Likely) |
| An Averostran theropod, possible Ceratosaur or Tetanuran. In 2024 was formally described, but referred to the older Blue Lias.[25] A paper non quoted on the description relocated it on the Charmouth mudstone.[26] | ||
Indeterminate | Near Lyme Regis | obtusum Zone, obtusum subzone |
| A theropod, possibly a basal member of Neotheropoda. Previously misidentified as a referred specimen of the pterosaur Dimorphodon macronyx.[27] NHMUK PV R36855 differs from those of all other Early Jurassic-Late Triassic, theropods in its combination of features, but bears no discernible autapomorphies.[26] It is similar to Tawa, Dracoraptor and Liliensternus, being considered an early branching neotheropod, with a size range similar to Coelophysis bauri, representing an individual of approximately 10 kg, approximately the same size as Coelophysis bauri.[26] | ||
Indeterminate | lower cliff face, Charmouth | obtusum Zone, obtusum subzone |
| A sauropodomorph, possibly a member of Massopoda. While it was not associated with BMNH 39496 or GSM 109560 and was not assigned to "Merosaurus" it was classified as coming from an indeterminate theropod. Mortimer found closer resemblance with pedal unguals of basal massopods like Blikanasaurus and Jingshanosaurus.[29] | ||
S. harrisonii |
| Black Ven Marl Member |
| A basal thyreophoran, and the type member of the family Scelidosauridae. One of the best known armored dinosaurs of the lower Jurassic, also the most complete found to date, with specimens preserving even the skin and dermal armour.[31] | ||
S. "sp. A" | Belemnite Marls at Seatown |
|
| |||
Cf. S. sp. | Foot of Black Ven, Charmouth | Asteroceras obtusum ammonoid zone, Black Ven Marl Member |
|
Fish
[edit]Numerous fish species are known from the Charmouth Mudstone and underlying Blue Lias, from such horizons as the "Fish Bed" of the Shales With Beef Member.
Fish of the Charmouth Mudstone Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Acrodus | A hybodont shark | |||||
Hybodus | A hybodont shark | |||||
Palidiplospinax | A synechodontiform shark | |||||
Squaloraja | S. tenuispina, S. polyspondyla | Closely related to modern chimaeras | ||||
Myriacanthus | M. paradoxus, M. granulatus | Black Ven Marl Member | A myriacanthid closely related to modern chimaeras | |||
Dorsetichthys | D. bechei | A stem-group teleost | ||||
"Coccolepis" | "C." liassicus | A coccolepidid fish, probably does not belong to the genus | ||||
Holophagus | H. gulo | A coelacanth | ||||
Chondrosteus | C. acipenseroides | A chondrosteid acipenseriform fish, related to sturgeon and paddlefish | ||||
Oxygnathus | O. ornatus | A palaeonisciform fish | ||||
Saurorhynchus | S. brevirostris, S. anningae | A member of Saurichthyiformes | ||||
Ptycholepis | P. gracilis, P. curtus | A palaeonisciform fish | ||||
Dapedium | Spp. | A dapediiform fish | ||||
Caturus | Spp. | An amiiform fish related to bowfins | ||||
Platysiagum | P. sclerocephalum | A platysiagid fish | ||||
Furo | F. orthostomus | A member of Ionoscopiformes within Halecomorphi |
Insects
[edit]Numerous species of insect are known from concretions, predominately in the Black Ven Marl Member.[32][33][34][35]
Insects of the Charmouth Mudstone Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Anglophlebia[36] | A. gigantea | Woodstones, Black Ven | Black Ven Marl Member | NHMUK In.51030, part and counterpart of partial forewing | Damsel-dragonfly | |
Archaeolepis[37] | A. mane | Birchi Nodules, Black Ven | Shales with Beef | Isolated wing | Among the oldest known lepidopterans | |
Archelcana[37] | A. durnovaria | Woodstones | Black Ven Marl Member | BMNH In 59162 | Elcanid orthopteran | |
Austaulius[38] | A. haustrum | Monmouth Beach, Lyme Regis | Black Ven Marl Member | NHMUK II 3103 | Necrotauliid caddisfly | |
Brevicula[37][39][40] | B. gradus, B. maculata | Flatstones, Woodstones;Monmouth Beach, Lyme Regis | Black Ven Marl Member | B. gradus: Holotype: NHMUK In.53993, female with ovipositor, B. maculata: Holotype: NHMUK II.3086, isolated Tegmina | Dermapterid earwig | |
Brochocoleus[37] | B. maculatus | Flatstones, Stonebarrow | Black Ven Marl Member | BMNH In 49577, 59133, 59361, 59154 | Ommatid beetle | |
Chrismooreia[41] | C. michaelbehei | Charmouth | Obtusum Zone | Part and counterpart of a mostly complete specimen | Asiopterid damsel-dragonfly | |
Dacryoderma[40] | B. teres | Charmouth-by-pass construction site | Obtusum Zone | Isolated tegmen | Dermapterid earwig | |
Daniilacheta[42] | D. aristovi | Monmouth Beach, Brooki bed | Shales with Beef Member | ZIN LD/Orth 19 | Protogryllidae cricket | |
Dorsettia[37] | D. laeta | Flatstones | Black Ven Marl Member | BMNH In 59375, a male hindwing fragment | Campterophlebiid damsel-dragonfly, genus also known from China | |
Durnovaria[37] | D. parallela | Flatstones | Black Ven Marl Member | BMNH In 59171 | Aerophasmatid, Stem group of Phasmatodea | |
Elaterina[37] | E. liassica | Lyme Regis | Black Ven Marl Member | BMNH In 59369 | Click beetle | |
Elaterophanes[39] | E. regius | Flatstones | Black Ven Marl Member | BMNH In 59385, 53952 | Click beetle | |
Eoptychoptera[37] | E. spectra | Flatstones | Black Ven Marl Member | BMNH In 49226 | Ptychopterid fly, formerly referred to the genus Prodocidia | |
Hagla[37] | H. cf. gracilis | Flatstones | Black Ven Marl Member | BMNH In 59167 | Haglidae hump-winged cricket | |
Holcoptera[36][37][39] | H. giebeli, H. alisonae; H. schlotheimi | Flatstones; Bed 75, Black Ven; Catherstone Lane; Monmouth Beach, Lyme Regis | Black Ven Marl Member | Complete specimen, isolated wings | Coptoclavid beetle | |
Hypsothemis[37] | H. fraseri | Flatstones | Black Ven Marl Member | BMNH In 59109, a hindwing | Campterophlebiid damsel-dragonfly, genus also known from China and Kazakhstan | |
Jurachorista[43] | J. bashkuevi | Monmouth Beach, Brooki bed | Shales with Beef Member | ISEA I-F/MP/4/1637/14 | Eomeropid scorpionfly | |
Lateophlebia[37] | L. anglicanopsis | Flatstones | Black Ven Marl Member | BMNH In 49573, 59376 | Campterophlebiid damsel-dragonfly | |
Lethonectes[44] | L. naucoroides | Flatstones | Black Ven Marl Member | BMNH In 49615, 51014(59383) | Belostomatidae giant water bug | |
Liassocorixa[44] | L. dorsetica | Flatstones | Black Ven Marl Member | BMNH In 59166 | Corixid bug | |
Liassocupes[36] | L. parvus | Flatstones | Black Ven Marl Member | BMNH In 64008, 49210 | Ommatid beetle | |
Liassophlebia[37] | L. pseudomagnifica | Stonebarrow | Black Ven Marl Member | NHMUK In.64000, partial hindwing | Liassophlebiid damsel-dragonfly | |
Locustopsis[37][42] | L. spectabilis | Flatstones; Monmouth Beach, Brooki bed | Black Ven Marl Member | ZIN LD/Orth 1, Orth 4, Orth 10, Orth 18, Orth 31 | Locustopsid grasshopper | |
Megarhyphus[45] | M. amberae | Monmouth Beach, Brooki bed | Shales with Beef Member | NHML In 28447 | Anisopodidae wood gnat | |
"Mesocixiella"[37] | "M." fennahi | Flatstones | Black Ven Marl Member | BMNH In 53942 | Fulgoridiid planthopper, does not belong to the genus Mesocixiella | |
Metaraphidia[37] | M. confusa | Charmouth (GSM) | Black Ven Marl Member | GSM 117552 | Snakefly | |
Micromacula[37] | M. gracilis | Flatstones, Birchi Nodules; Flatstones, Black Ven | Shales with Beef; Black Ven Marl Member | BMNH In 49230 | Regiatid orthopteran | |
Mimemala[37] | M. giganteum | Flatstones | Black Ven Marl Member | BMNH In 51026 | Schizocoleid beetle | |
Nannoblattina[37] | N. petulantia | Flatstones | Black Ven Marl Member | BMNH In 53929, 51004 | Mesoblattinid cockroach | |
Nannotanyderus[46] | N. oliviae | Monmouth Beach, Brooki bed | Shales with Beef Member | ISEA I-F/MP/2/1621/13, 12 | Tanyderid fly | |
Neomeridium[44] | N. trifurcum | Woodstones | Black Ven Marl Member | BMNH In 59151, 51039, 53901, 64014 | Pachymeridiid Lygaeoid bug | |
Oligophryne[47] | O. britannica | Stonebarrow Hill | Black Ven Marl Member | Coll. Ansorge fly No 1 | Oligophrynidae fly | |
Omma[37] | O. liassicum | Flatstones | Black Ven Marl Member | BMNH 59132 | Ommatid beetle, genus extant | |
Orthophlebia[37] | O. capillata | Flatstones | Black Ven Marl Member | BMNH In 53924, an exoskeleton (wings & thorax) | Orthophlebiid scorpionfly | |
Orichalcum[37] | O. ornatum | Flatstones, Birchi Nodules | Shales with Beef | BMNH In 53983 | Triassomanteidae grasshopper | |
Paraprosbole[37] | P. rotruda | Flatstones | Black Ven Marl Member | BMNH In 59374, 48162, 64395 | Tettigarctid cicada | |
Propreocoris[44] | P. maculatus | Woodstones | Black Ven Marl Member | BMNH In 59152 | Stem-group to Ochteridae and Gelastocoridae | |
Protogryllus[37] | P. (Archaegryllodes) magnus | Flatstones | Black Ven Marl Member | BMNH In 44003, 51016, 51022, 59373 | Protogryllidae cricket | |
Protohagla[36][37] | P. langi | Flatstones | Black Ven Marl Member | BMNH In 59018 | Haglid cricket | |
Protomyrmeleon[41] | P. cf. brunonis | Charmouth burden pile | Black Ven Marl Member | AR 428 | Protomyrmeleontidae winged insect | |
Protorthophlebia[37] | P. latipennis | Flatstones | Black Ven Marl Member | BMNH In 49575, 53954, 53963, 59120, 59365, 59371, 59384 | Protorthophlebiid scorpionfly | |
Priscaenigma[37] | P. obtusa | Flatstones | Black Ven Marl Member | BMNH In 53898 | Snakefly | |
Pseudopolycentropus[36] | P. triangularis | Woodstones | Black Ven Marl Member | BMNH In 53915 | Pseudopolycentropodid scorpionfly | |
Pterocimex[44] | P. jacksoni | Woodstones | Black Ven Marl Member | BMNH in 53909, 49589, 51048, 59359 | Nepomorphan | |
Regiata[37] | R. scutra | Flatstones | Black Ven Marl Member | BMNH In 64027 | Regiatid orthopteran | |
Rhipidoblattina[37] | R. ssp. | Flatstones | Black Ven Marl Member | BMNH In 49212, 49249, 53909, 53920 | Caloblattinidae cockroach | |
Rossiphlebia[37] | R. jacksoni | Flatstones | Black Ven Marl Member | NHMUK In.53999, part and counterpart of a partial hindwing | Liassophlebiid damsel-dragonfly | |
Tarsabedus[44] | T. menkei | 0.6 km east Charmouth | Black Ven Marl Member | BRSMG Cc 688 | Belostomatidae giant water bug | |
Tersus[37] | T. crowsoni | Flatstones | Black Ven Marl Member | BMNH In 53949, 59038 | Schizophorid beetle |
Plants
[edit]Genus | Species | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|
Ctenozamites[48] |
|
| Isolated shoots | A "Seed Fern" of the group with Corystospermales | |
Cycadites[48] |
|
| Isolated shoots | A possible Cycadophyte leaf, related with Cycadales. Alternatively, it can be of Bennettite origin | |
Cycadeoidea[48] |
|
| Stems | Bennettite Trunks of the family Cycadeoidaceae | |
Cycadopteris[48] |
|
| Isolated shoots | A "Seed Fern" of the group with Corystospermales | |
Otozamites[49] |
|
| Isolated shoots | Bennettite leafs of the family Williamsoniaceae | |
Pagiophyllum[49] |
|
| Isolated shoots | Conifer shoots of the family Hirmeriellaceae or Araucariaceae |
See also
[edit]References
[edit]- ^ British Geological Survey. "Charmouth Mudstone Formation". BGS Lexicon of Named Rock Units. Retrieved 2018-03-08.
- ^ Gallois, R.W. (2008). The lithostratigraphy of the Shales-with-Beef Member of the Charmouth Mudstone Formation, Lower Jurassic (PDF). Ussher Society. OCLC 703253954.
- ^ "Black Ven Marl Member". The BGS Lexicon of Named Rock Units. British Geological Survey.
- ^ "Belemnite Marl Member". The BGS Lexicon of Named Rock Units. British Geological Survey.
- ^ "Green Ammonite Member". The BGS Lexicon of Named Rock Units. British Geological Survey.
- ^ Owen (1865). "Monograph of the Fossil Reptilia of the Liassic Formations. Part First. Sauropterygia". Monographs of the Palaeontographical Society. 17 (75): 1–40. Bibcode:1865MPalS..17....1O. doi:10.1080/02693445.1865.12027939. ISSN 0269-3445.
- ^ Sollas, W. J. (1881). "On a new Species of Plesiosaurus (P. Conybeari) from the Lower Lias of Charmouth; with Observations on P. megacephalus, Stutchbury, and P. brachycephalus, Owen". Quarterly Journal of the Geological Society. 37 (1–4): 440–481. doi:10.1144/gsl.jgs.1881.037.01-04.42. ISSN 0370-291X.
- ^ Storrs, G. W. (1995). "A juvenile specimen of ? Plesiosaurus sp. from the Lias (Lower Jurassic, Pliensbachian) near Charmouth, Dorset, England". Proceedings of the Dorsal Archaeological and Natural History Society. 116 (2): 71–76.
- ^ CONYBEARE, W. D. (1824). "XXI.—On the Discovery of an almost perfect Skeleton of the Plesiosaurus". Transactions of the Geological Society of London. 1 (2): 381–389. doi:10.1144/transgslb.1.2.381. ISSN 2042-5295.
- ^ Smith, Adam S.; Araújo, Ricardo (2017-07-21). "Thaumatodracon wiedenrothi, a morphometrically and stratigraphically intermediate new rhomaleosaurid plesiosaurian from the Lower Jurassic (Sinemurian) of Lyme Regis". Palaeontographica Abteilung A. 308 (4–6): 89–125. Bibcode:2017PalAA.308...89S. doi:10.1127/pala/308/2017/89. ISSN 0375-0442.
- ^ Lomax, Dean R.; Massare, Judy A. (2015-03-04). "A new species of Ichthyosaurus from the Lower Jurassic of West Dorset, England, U.K." Journal of Vertebrate Paleontology. 35 (2): e903260. Bibcode:2015JVPal..35E3260L. doi:10.1080/02724634.2014.903260. ISSN 0272-4634. S2CID 85745787.
- ^ Bennett, S.P.; Barrett, P.M.; Collinson, M.E.; Moore-Fay, S.; Davis, P.G.; Palmer, C.P. (January 2012). "A new specimen of Ichthyosaurus communis from Dorset, UK, and its bearing on the stratigraphical range of the species". Proceedings of the Geologists' Association. 123 (1): 146–154. Bibcode:2012PrGA..123..146B. doi:10.1016/j.pgeola.2011.07.001.
- ^ McGowan, Christopher; Milner, Angela C. (October 1999). "A new Pliensbachian ichthyosaur from Dorset, England". Palaeontology. 42 (5): 761–768. Bibcode:1999Palgy..42..761M. doi:10.1111/1475-4983.00096. ISSN 0031-0239.
- ^ Lomax, Dean R.; Massare, Judy A. (December 2018). "A forefin of Leptonectes solei from the Lower Jurassic (Pliensbachian) of Dorset, UK". Proceedings of the Geologists' Association. 129 (6): 770–773. Bibcode:2018PrGA..129..770L. doi:10.1016/j.pgeola.2018.07.005. S2CID 134397018.
- ^ "PBDB".
- ^ Young, Mark T; Dufeau, David; Bowman, Charlotte; Cowgill, Thomas; Schwab, Julia A; Witmer, Lawrence M; Herrera, Yanina; Katsamenis, Orestis L; Steel, Lorna (2024). "Thalattosuchian crocodylomorphs from the Sinemurian (Early Jurassic) of the UK". academic.oup.com. Zoological Journal of the Linnean Society. Retrieved 2024-07-15.
- ^ Wilberg, E. W.; Godoy, P. L.; Griffiths, E. F.; Turner, A. H.; Benson, R. B. J. (2023). "A new early diverging thalattosuchian (Crocodylomorpha) from the Early Jurassic (Pliensbachian) of Dorset, U.K. and implications for the origin and evolution of the group". Journal of Vertebrate Paleontology. 42 (3). e2161909. doi:10.1080/02724634.2022.2161909. S2CID 256149424.
- ^ Sangster, Sarah (2021-09-02). "THE OSTEOLOGY OF DIMORPHODON MACRONYX, A NON-PTERODACTYLOID PTEROSAUR FROM THE LOWER JURASSIC OF DORSET, ENGLAND". Monographs of the Palaeontographical Society. 175 (661): 1–48. Bibcode:2021MPalS.175....1S. doi:10.1080/02693445.2021.2037868. ISSN 0269-3445.
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